The bear then turned on Huguenard, who was also killed in the incident.
[caption id="attachment_372271" align="alignnone" width="1428"]
Treadwell was 46 when he was mauled to death by a grizzly bear (YouTube)[/caption]
The attack happened just hours before the couple were due to fly back home by seaplane for the winter.
Willy Fulton - who was the couple's air taxi pilot for the flight home - knew immediately what had happened when he landed at Katmai National Park.
Usually, the couple would be waiting on the shore to be picked up by Fulton, but on this occasion there was an eerie silence and the "meanest looking bear" standing on top of a pile of human remains.
The couple's tent was torn apart alongside their evening meal that was unopened.
A ranger spotted fingers and an arm in a 3ft-high mound of grass, mud and twigs, alongside other remains.
They also found nearby what was left of Treadwell's head, which was connected to a small piece of his spine.
The couple always filmed the bear interactions they had, and this occasion was no different.
In the panic as the bear attacked, they hadn't removed the camera lens.
But the equipment still recorded the audio of the horrific attack, and recorded the final moments of Treadwell's life.
Unverified audio has been shared on YouTube, which reportedly matches up with the coroner's report into the tragedy.
https://www.youtube.com/watch?v=hXyQAtXJ4II
The bear was eventually killed as part of rangers' attempts to retrieve the couple's remains.
In the years that followed, a documentary about the tragic attack was made by Werner Herzog titled
Grizzly Man.
The film was pieced together from Timothy's actual video footage of the bears, and examined the bear enthusiast's love of heading to the Alaskan reserve every year.
https://www.youtube.com/watch?v=uWA7GtDmNFU&t=34s
The biography was critically acclaimed, achieving a 92 percent Certified Fresh rating on Rotten Tomatoes.